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In B. lichenformis a few amino acids are removed from the C-terminal end of the

ID: 188176 • Letter: I

Question

In B. lichenformis a few amino acids are removed from the C-terminal end of the beta-lactamase enzyme after it is synthesized. The sequence of the original C-terminus can be deduced by comparing it to a mutant in which the reading frame is shifted by insertion or deletion of a nucleotide. the amino acid sequence of the purified wild-type enzyme and the frameshift mutant from amino acid 263 to the C-terminus are given below.

Wild-type: N M N G K

Mutant: N M I W Q I C V M K D

1. What was the mutational event that gave rise to the frameshift mutation?

2. Deduce the number of amino acids in the synthesized (pro-enzyme) from the wild-type enzyme and, as far as possible, the actual sequence of the wild-type enzyme.

Explanation / Answer

wild type - NMNGK

CODON SEQUENCE - AAU AUG AAU GGC AAA

MUTANT - NMIWQCVMKD

CODON SEQUENCE - AAU AUG AUA UGG CAA AUA UGU GUA AUG AAA GAU

in the mutant insertion of U nucleotide shifted the frame. if the U (underlined and bold) was not inserted then the wild-type sequence will be -

NMNGKYV#(STOP CODON) = 269 AMINO ACID LONG WILD TYPE PROTEIN.

CODON SEQUENCE - AAU AUG AAU GGC AAA UAU GUG UAA